Rhoptromeris haywardi (Blanchard, 1947) is not Rhoptromeris Förster, 1869: new combination in Leptopilina Förster, 1869 (Hymenoptera: Cynipoidea: Figitidae: Eucoilinae)

. Rhoptromeris haywardi (Blanchard, 1947) is reviewed and the new combination Leptopilina haywardi (Blanchard, 1947) is proposed. Leptopilina haywardi comb. nov . is redescribed and illustrated. Leptopilina Förster, 1869 is cited for the first time for Uruguay and Tephritidae (Diptera) is reported for the first time as its host.

Rhoptromeris haywardi was originally described in Eucoila Westwood, 1833 by Blanchard in 1947, which designates two specimens as cotypes these correspond to syntypes according to article 73.2.1 of the ICZN (ICZN 1999).Subsequently, De Santis (1967) made the transfer to Rhoptromeris.This species is characterized by having pronotal plate with incomplete lateral bridges and open lateral fovea; characters that indicate that it is not a member of Trichoplastini and, therefore, it is not a Rhoptromeris.In addition, R. haywardi is only species of Rhoptromeris cited for the Neotropical region, with reported presence in Argentina and Uruguay; it is also the only one that has Tephritidae (Diptera) reported as a host (Turica & Mallo 1961;De Santis 1967;Garcia et al. 2020).So to which genus does this species belong?
We compared R. haywardi with Ganaspis Förster, 1869, Trybliographa Förster, 1869, and Leptopilina Förster 1869, because many of the species included in these genera behave as parasitoids of frugivorous dipterous of families Drosophilidae and Tephritidae.(Blanchard, 1947) that is, in this way, reported for the first time for Uruguay.
There are 41 species described species of Leptopilina worldwide (Buffington et al. 2020), and those that have a known host mainly attack frugivorous drosophila flies (Diptera: Drosophilidae) (Lue et al. 2016).
Leptopilina haywardi comb.nov., is similar to Leptopilina heterotoma (Thomson, 1862) and Leptopilina pacifica Novkovic & Kimura, 2011, by a large scutellar plate rhomboid-shaped (a true rhombus-shaped in L. heterotoma and teardrop-shaped in L. pacifica).It differs from both species by presenting the following combination of characters: head, meso and metasoma reddish to light brown (black or dark brown in L. heterotoma and reddish brown to dark brown in L. pacifica); female antenna semiclavate (clavate in L. heterotoma and L. pacifica), with placoidal sensillas present on F5-F11 (present on F6-F11 in L. heterotoma and on F5 or F7-F11 in L. pacifica); foveas at base of scutellum oval (semi-squares in L. heterotoma and L. pacifica); dorsal surface of scutellum areolate (puntate-reticulate or reticulaterugose in L. heterotoma and reticulate in L. pacifica); metapleura with ridges 1 and 2 fully developed and ridge 3 extending beyond half of metapleura (3 ridges fully developed in L. heterotoma and ridges 1 and 2 fully development and ridge 3 short in L. pacifica); middle area of propodeum with scattered hair, laterals densely haired (entire surface

Taxonomic Authorities
Rhoptromeris haywardi presents the following combination of characters: head narrows ventrally, triangle-shaped in frontal view (semicircular in frontal view in Ganaspis and usually rounded in frontal view in Trybliographa); dorsal margin of pronotal plate straight (slightly emarginated in Ganaspis and never straight in Trybliographa); laterals bars of scutellum short (long in Trybliographa and in short or long in Ganaspis); lateral depression of scutellum undefined (well defined in Ganaspis and defined in Trybliographa); marginal cell of forewing closed (open or partiality open in Trybliographa and, in Ganaspis, can also be closed); metapleural ridges well defined (metapleura smooth in Ganaspis; ridges 1 and 3 present, ridge 2 absent in Trybliographa); petiole enlarged posteriorly and the broad posterior rim with sculptural patterns (without posterior rim in Ganaspis and Trybliographa); base of syntergum with a hairy ring more or less reduced only ventrally, interrupted dorsally (dense hair ring and usually complete dorsally in Ganaspis; always complete dorsally in Trybliographa).This combination of characters corresponds to Leptopilina.Therefore, we propose the new combination Leptopilina haywardi 2 females, 10-XI-1936, s/Anastrepha, Ins.Pat.Vegetal, INTA (MLP).Additional material: ARGENTINA.Entre Ríos, Concordia.5 females, VI-1936, s/Anastrepha fraterculus (MLP).